I think, my brain is fried right now with other things, this is how it goes.
They all signed WADA, UCI is under the IOC umbrella. USAC is a NGB under the USOC which falls under the IOC.
Bottom of page 3 and and page 4 is where the answer lies in the article.
USAC falls under USOC/IOC and is the licensing body.
"USADA has been delegated the anti-doping responsibilities that were previously shared by the USOC and the more than forty (40+) NGBs in the United States"
"A significant achievement of WADA was the drafting and adoption in 2004 of the World Anti-Doping Code (the “Code”), a consistent body of rules which inform the anti-doping rules adopted by each of the international federations (IFs) responsible for governing each individual Olympic sport." (Cycling is an Olympic sport regardless if you ride in the Olympics.)
"In addition to WADA and national anti-doping organizations (NADOs) such as USADA, most of the IFs also retain responsibility for anti-doping matters within their sport. Thus, for example, WADA, USADA and the International Association of Athletics Federations (IAAF)(exchange with USA Cycling), the international federation which governs the sport of track and field, have overlapping authority to conduct drug testing on elite U.S. track and field athletes who compete in international competition. This overlapping authority helps to ensure a level playing field."
"When an athlete becomes a member of a NGB they agree to submit their
eligibility disputes, including disputes concerning anti-doping rule violations, to arbitration." USA Cycling is a NGB, Lance obtained a license yearly to race internationally through USA Cycling. UCI does not give individual rider licenses.