We must not forget that they are not really teammates. The Russians have different training gropus who are in fierce competition with each other. So it does not come as a suprise that they do not really cooperate. I think all in all they benifit of that competitive environment, though.
The idea of team tactics in individual races is a bit strange anyway. This is not cycling were they are all getting paid by their teamsports. It is an individual sport and in my opinion the relay is the only time there should be team tactics. In all the other racers everyone should race for his own success. No matter the nation.
Yes and I think those groups only see each other when there is a joint testing week in September. Then it's sometimes for the pre-world cup warmup races and of course they race the world cups/world champs/olympics, but even then, you can see that those groups don't really travel together, just stay at the same hotel, as part of the national team.
Ustiugov was diplomatic and seemed content that they went 1-2, but he was also disappointed that he couldn't shake Bolshunov. Last year Bolshunov was part of the Russian led group that skied away from Ustiugov in the Val di Fiemme 15km, and he lost the tour that day. He had too much to overcome on the final day. I am sure that looms in his head as well.