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The quoted section that I posted is from a 'copy & paste' from an online pdf of the article - and the word in question IS "by" in the article.
The pdf is at -
http://www.google.com/url?sa=t&sour...sg=AFQjCNG4mgIRO33yxt3RXiPVNBHxoYLMnA&cad=rja
and the quoted section is the last paragraph on page 106.
Also, I do not think the word 'by' is a typo - use of the word 'be' in that sentence is very clumsy and inconsistent with the proper English in the rest of the article.
Regarding the pedaling style of the participants, this is from pages 99-100 -
"Interestingly, the subjects of
group 1 produced the larger propulsive torques by
creating significantly (P < 0.05) larger forces in the
vertical direction on the pedal during the downstroke
(peak Fy in Table 6 and Fig. 6b) and by not attempting
to pull up on the pedal during the upstroke (near zero
values for Fy in Fig. 6b between 0 = 240° and 0 = 340°).
The pattern of horizontal force application to the pedal
was more similar in the two groups (Fig. 6c).
These data indicated that the "elite" cyclists in group
1 adopted a pedaling strategy of applying large vertical
forces to the pedal during the downstroke in an attempt
to maximize the amount of propulsive torque produced
during this interval. Figure 5 indicates that group 2
used a similar strategy of generating propulsive torque
during the downstroke, with two small differences.
Group 2 applied smaller forces during the downstroke
and sufficient upward force during recovery to generate
small propulsive torques."
The participants for the study WERE NOT chosen based on their pedaling style, and there was no attempt made in the study to have them change their style.
I suggest that anyone interested in this article takes the time & effort to read it completely and carefully to understand what it DOES say. 
Jay Kosta
Endwell NY USA