Optimal cadence selection during cycling
"Cadence or pedal rate is widely accepted as an important factor influencing economy of motion, power output, perceived exertion and the development of fatigue during cycling."
I especially like Figure 1a which shows how pedaling technique changes with power in that the higher the power, not only does the rider push harder on the down stroke but the rider does less negative work (and more positive work) on the upstroke. And, also, that it changes with cadence showing it is easier to get the rising foot out of the way on the upstroke when pedal speed is reduced. (not that the "just push harder crowd" cares).
But, here is the meat of the paper as it regards oxygen consumption. "Numerous studies have examined the influence of pedalling frequency on the efficiency and economy of cycling 6, 27, 44, 46, 48, 71, 72. Generally, when cycling at constant power outputs, lower cadences have been found to result in reduced oxygen cost (i.e. improved gross efficiency) compared with higher cadences 6, 15, 27. Improved efficiency of cycling observed at lower pedalling rates is likely to be dictated by the relationship between muscle shortening velocity and the efficiency of muscle contractions (percent Type I and Type II active fibres). For instance, under in vitro conditions, it has been observed that the efficiency of skeletal muscle contractions is augmented with increasing speed of contraction, until a maximum is reached (i.e. an economically optimal shortening velocity) 35. The most economical cadence appears to be extremely low (~50-60rpm) when cycling at low power outputs (?W), but increases to approximately 80-100rpm with increasing workloads (~350W) 26, 44, 58. The cause of the rise in the economically optimal cadence is unclear, but is again likely to be due to the power-velocity relationship of muscle contraction and the additional recruitment of fast twitch muscle fibres with increases in exercise intensity."